Ok I guess I kind of killed this thread with that last post, rofl .Let me see if I can be a little better at explaining it.
Ok in a sediment filter (ie a dsb/ssb/cc/lr) things do not quite happen the way most folks understand it. This is one of the main problems with maintaining one. The nitrifing/denitrifing/reducing/oxidising is all bacterial driven, nothing else, just bacteria. Forget about sand bed critters, forget about cukes, or blennies or what have you, its all about the bacteria.
Now in the sediment system folks tend to follow the zonal thing, as in ammonia/nitrite bacteria live in the aerobic zone and denitrifing bacteria live in the anaerobic zone. Ok you need to forget that, although it is true it is but one branch of a multi-armed process. You need to look at a sediment bed as an ever moveing continious state of flux. Every area on and in the substraight is always changing in all conditions (ie the enviroment is changing). In the bed thier are a many many differing strains of bacteria. they will come to the for front based on the conditions at hand. If the enviroment is skewed to thier liking they will dominate.
Ok I am getting to deep again, lol. Nitrification can happen at any level in your bed, so can denitrification, it all depends on the enviroment created, in the wild alot occurs in the water column (but that is another story, heheh). If certain conditions favor one bacterial strain ovver another it will dominate, its byproduct will then again allow a different bacteria to dominate as a result of its activities. these byproducts can make it or break it as far as if you will get any exportation or if you will be plagued by algae/cyano and so on so it pertains.
An example. Jeaubert plenum in Monico. at first it was closed, testing observed that ammonia and ammonium was present at almost all levels, bu not in the tank. With the presence of ammonia in the bed it would not allow denitrifing bacteria to tanke nitrate to gas, (ammmonia inhibits the first enzyme in denitrification) .in stead other strains took the nitrate and converted it into ammonium and put it back into cycle just above nitrite. Thus this filter lost its ability to export nitrogen products and became an ever growing nitrogen sink. .
Ok lets see if that sparks anyone, lol or if its doomed to die a slow death.
Mike